1982-1987 PhD, King's College, University of London
1987-1990 Post-doc, Case Western Reserve University
1990-1993 Post-doc, Temple University
1993-2000 Assistant Professor, Department of Biological Sciences, Ohio University
2000-2001 Associate Professor, Department of Biological Sciences, Ohio University
2001-2010 Associate Professor, Department of Neuroscience, Ohio State University
2010-present Professor, Department of Neuroscience, Ohio State University
2015-present Co-Director, Graduate program in Molecular, Cellular and Developmental Biology, Ohio State University